Cynodon dactylon

Family:POACEAE
Species:Cynodon dactylon (L.) Pers.
Common Name:Bermuda Grass; Scutch Grass
Habitat:**
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Perennial
Category:Vascular
USDA Symbol:**
Plant Notes:Bermuda Grass is an introduced herbaceous perennial in the Grass family (Poaceae). It occurs widely in tropical and subtropical areas around the world, and its native range is uncertain. It most likely originated in tropical Africa, the Mediterranean area, or India/Bangladesh. The first record of it being imported into the United States is from Savannah Georgia in 1751 by Governor Henry Ellis. It was likely cultivated earlier by the Spanish in Florida. Bermuda Grass occurs throughout Alabama. It is cultivated in lawns and on sports fields, planted for erosion control, and is used as a pasture grass for hay production. It escapes to roadsides, field edges, vacant lots, and other disturbed sites. Bermuda Grass is a warm season carpet grass with long scaly rhizomes and thin stolons. It is highly variable in growth form and many cultivars and hybrids have been developed. Common turf varieties such as `Bradley', `Sunturf', `Tiffine', and `Tiflawn' are hybrids between Bermuda Grass and African Bermuda Grass (C. transvaalensis Burtt-Davy). Bermuda Grass culms are erect and vary from 1-20 inches in height. The sheaths are glabrous with long white hairs at the collar. The leaves are ½ - 2 ½ inches in length, linear in outline, flat, and glabrous or pubescent on the lower surface. Flowers are produced in panicles. The panicles usually have 4-6 branches in a single whorl. The fruit is a grain. The leaves of Bermuda Grass are rich in Vitamin C and are crushed and made into a tea in India. Bermuda Grass has also been used as a home remedy for heartburn or indigestion, and as a charm to ward off sorcery. It is a common weed of agricultural fields and can be difficult to eradicate once established.—A. Diamond.
Taxonomic Notes:**
Status:Not Native
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyPOACEAE - Grass family
Genus Cynodon
Species Cynodon dactylon (L.) Pers. - Bermuda Grass; Scutch Grass

Citation

Citation Cynodon dactylon (Linnaeus) Persoon, Syn. Pl. 1: 85. 1805.
Basionym: Panicum dactylon L. 1753.
Type: PORTUGAL: Without data (lectotpye: LINN 80.35). Lectotypified by Clayton & J. R. Harlan, Kew Bull. 24: 185-189. 1970.

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Chloris cynodon Chloris cynodon Trinius, Gram. Unifl. Sesquifl. 229. 1824, nom. illegit.Panicum dactylon Linnaeus 1753. 
Cynodon aristiglumis Cynodon aristiglumis Caro & E. A. Sánchez, Kurtziana 5: 236, f. 11. 1969. ARGENTINA: Río Negro: Dpto. Avellaneda: Isla Choele-Choel, Brazo Sud del Río Negro-Sifón, 14 Feb 1948, Boelcke & Serrano 3171 (holotype: BAA).
Cynodon aristulatus Cynodon aristulatus Caro & E. A. Sánchez, Darwiniana 17: 514, f. 2. 1972 PERU: San Martín: Tingo María near Huallaga River, 30 Oct 1949-19 Feb 1951, Allard 22027 (holotype: US).
Cynodon dactylon var. elegans Cynodon dactylon (Linnaeus) Persoon, var. elegans Rendle, in Hiern, Cat. Afr. Pl. 2: 221. 1899. ANGOLA: Mocamedes, Cavalheiros to mouth of R. Bero, s.d., Welwitsch 2290 (holotype: ?; isotype: K).
Cynodon dactylon var. glabratus Cynodon dactylon (Linnaeus) Persoon, var. glabratus (Steudel) Chiovenda, Nuovo Giorn. Bot. Ital., ser. 2. 26: 82. 1919.Cynodon glabratus Steudel 1854. 
Cynodon dactylon subsp. glabratus Cynodon dactylon (Linnaeus) Persoon, subsp. glabratus (Steudel) Chevalier, Rev. Int. Bot. Appl. Agric. Trop. 27: 282. 1947.Cynodon glabratus Steudel 1854. 
Cynodon erectus Cynodon erectus J. Presl, in C. Presl, Reliq. Haenk. 1: 290. 1830. PERU [mis-cited as also in Mexico]: Without data, Haenke s.n. (holotype: PR).
Cynodon glabratus Cynodon glabratus Steudel, Syn. Pl. Glumac. 1: 212. 1854. ETHIOPIA
Cynodon pascuus Cynodon pascuus Nees von Esenbeck, in Martius, Fl. Bras. Enum. Pl. 2(1): 425. 1829. BRAZIL: Piauí: Cachian and vicinity, s.d., Martius s.n. (holotype: M; isotype: US).
Cynodon polevansii Cynodon polevansii Stent, Bothalia 2: 283. 1927. SOUTH AFRICA: Transvaal: Berber's Pen, s.d., Pole-Evans 334 (holotype: PRE).
Cynodon tenuis Cynodon tenuis Trinius, in Sprengel, Neue Entd. 2: 63. 1821.  
Cynodon umbellatus Cynodon umbellatus (Lamarck) Caro, Dominguezia 6: 16. 1983Paspalum umbellatum Lamarck 1791. 
Cynosurus uniflorus Cynosurus uniflorus Walter, Fl. Carol. 82. 1788. SOUTH CAROLINA
Dactilon officinale Dactilon officinale Villars, Hist. Pl. Dauphine 2: 69. 1787, nom. illegit.Panicum dactylon Linnaeus 1753. 
Digitaria dactylon Digitaria dactylon (Linnaeus) Scopoli, Fl. Carn., ed. 2. 1: 52. 1772.Panicum dactylon Linnaeus 1753. 
Digitaria glumaepatula Digitaria glumaepatula (Steudel) Miquel, Fl. Ned. Ind. 3: 439. 1857.Panicum glumaepatulum Steudel 1853. 
Digitaria littoralis Digitaria littoralis Salisbury, Prodr. Stirp. Chap. Allerton 19. 1796, nom. illegit.Panicum dactylon Linnaeus 1753. 
Digitaria stolonifera Digitaria stolonifera Schrader, Fl. Germ. 165. 1806, nom. illegit.Panicum dactylon Linnaeus 1753. 
Fibichia dactylon Fibichia dactylon (Linnaeus) Beck, Fl. Bosnien 9: 436. 1904.Panicum dactylon Linnaeus 1753. 
Fibichia umbellata Fibichia umbellata Koeler, Descr. Gramin. 308. 1802, nom. illegit.Panicum dactylon Linnaeus 1753. 
Milium dactylon Milium dactylon (Linnaeus) Moench, Methodus 67. 1802.Panicum dactylon Linnaeus 1753. 
Panicum dactylon Panicum dactylon Linnaeus, Sp. Pl. 58. 1753. PORTUGAL: Without data (lectotpye: LINN 80.35). Lectotypified by Clayton & J. R. Harlan, Kew Bull. 24: 185-189. 1970.
Panicum glumaepatulum Panicum glumaepatulum Steudel, Syn. Pl. Glumac. 1: 41. 1853. PHILIPPINES: Luzon: Without data, Cuming 550 (holotype: P; isotypes: L, MO).
Paspalum dactylon Paspalum dactylon (Linnaeus) Lamarck, Tabl. Encycl. 1: 176. 1791.Panicum dactylon Linnaeus 1753. 
Paspalum umbellatum Paspalum umbellatum Lamarck, Tabl. Encycl. 1: 177. 1791. MAURITIUS: Without data, Commerson s.n. (holotype: P; isotype: US).
Capriola dactylon Capriola dactylon (Linnaeus) Kuntze, Revis. Gen. Pl. 2: 764. 1891.Panicum dactylon Linnaeus 1753. 

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Bibb, Blount, Bullock, Butler, Calhoun, Cherokee, Choctaw, Clay, Cleburne, Coffee, Conecuh, Coosa, Covington, Crenshaw, Cullman, Dale, DeKalb, Escambia, Etowah, Fayette, Franklin, Jackson, Jefferson, Lee, Madison, Marengo, Marion, Marshall, Mobile, Montgomery, Morgan, Pickens, Pike, Randolph, Russell, Shelby, St. Clair, Talladega, Tallapoosa, Tuscaloosa, Walker, Winston

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: AUA, JSU, SAMF, TROY, UNA, UWAL

Range of years during which specimens were collected: 1952 - 2023

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
JSU108277 Lee 24 Jul 1973 Johnson, Kenneth G.
128
JSU
Cynodon dactylon (Linnaeus) Persoon
JSU108349 Winston 07 Jun 1975 Whetstone, R. David
4492
JSU
Cynodon dactylon (Linnaeus) Persoon
JSU108274 Jefferson 13 Jul 1984 Barber, J. Patrick
821
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015991 Lee 22 May 1970 Evans, R.W.
153
AUA
Cynodon Dactylon (L.) Pers.
JSU108345 St. Clair 13 Jul 1969 Ellard, Jerry
s.n.
JSU
Cynodon dactylon (Linnaeus) Persoon
SAMF0001703 Mobile 13 Apr 2013 Davenport, Lawrence J.
5429
SAMF
Cynodon dactylon (Linnaeus) Persoon
JSU156468 Etowah 27 May 1994 Hodge, Lesley M.
1327
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015984 Butler 13 Aug 2005 Diamond, Alvin R.
15633
AUA
Cynodon Dactylon (L.) Pers.
SAMF0007748 Mobile 25 Apr 2015 McPheeters, Kendall
30
SAMF
Cynodon dactylon (Linnaeus) Persoon
JSU108272 Jackson 26 May 1999 Brodeur, Kristin R.
1842
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000033639 Barbour 01 Oct 2011 Morrison, Derek
137
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108270 Etowah 18 Jul 1969 Smith, Doris
s.n.
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000011822 Butler 13 Aug 2005 Diamond, Alvin R.
15634
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108278 Marshall 24 May 1992 Spaulding, Daniel D.
1863
JSU
Cynodon dactylon (Linnaeus) Persoon
UNA00003688 Mobile 26 Aug 1965 Deramus, Rebecca
D731
UNA
Cynodon dactylon (L.) Pers. var. dactylon
JSU108275 Jefferson 28 May 1984 Barber, J. Patrick
398
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000016001 Russell 1 Jul 2002 Gil, Glenda
2001-256
AUA
Cynodon Dactylon (L.) Pers.
JSU108343 Shelby 11 Oct 1997 Spaulding, Daniel D.
10208
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015986 Coffee 23 Jul 1989 Diamond, Alvin R.
6068
AUA
Cynodon Dactylon (L.) Pers.
JSU108346 Shelby 10 Sep 1982 Whetstone, R. David
12262
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000038384 Morgan 11 Jun 2017 England, J. Kevin
7983
AUA
Cynodon Dactylon (L.) Pers.
SAMF0008206 Butler 13 Aug 2005 Diamond. Alvin R.
15634
SAMF
Cynodon dactylon (Linnaeus) Persoon
AUA000016000 Pike 7 May 1994 Diamond, Alvin R.
9135
AUA
Cynodon Dactylon (L.) Pers.
JSU108341 Randolph 15 Jul 1988 Nixon, Christopher F.
4038
JSU
Cynodon dactylon (Linnaeus) Persoon
JSU108266 Cullman 20 Jun 1975 Whetstone, R. David
4881
JSU
Cynodon dactylon (Linnaeus) Persoon
JSU108261 Calhoun 27 May 1982 Whetstone, R. David
11196
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015996 Lee 1 Nov 1959 Banks, Donald J.
AUA
Cynodon Dactylon (L.) Pers.
UNA00064057 Shelby 10 Sep 1982 Whetstone, R.D.
12262
UNA
Cynodon dactylon (L.) Pers.
TROY000028200 Talladega 11 Oct 2008 Berry, Brittanni
10
TROY
Cynodon dactylon (Linnaeus) Persoon
AUA000015999 Lee 26 Aug 1934 Strangler, W.J.
AUA
Cynodon Dactylon (L.) Pers.
UNA00003634 Tuscaloosa 28 Apr 1955 Farmer, Joe A.
s.n.
UNA
Cynodon dactylon (L.) Pers. var. dactylon
AUA000015985 Cherokee 20 May 1972 Hale, A.R.
155
AUA
Cynodon Dactylon (L.) Pers.
TROY000011823 Butler 17 Aug 2004 Diamond, Alvin R.
14613
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108269 Etowah 10 Aug 1974 Atkinson, Tim A.
606
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000011834 Pike 14 May 1997 Evers, Astrid
134
TROY
Cynodon dactylon (Linnaeus) Persoon
AUA000015994 Lee 21 Nov 1962 Rogers, Ken E.
AUA
Cynodon Dactylon (L.) Pers.
UNA00003668 Tuscaloosa _ Jun 1955 Robinson, Flynt
s.n.
UNA
Cynodon dactylon (L.) Pers. var. dactylon
AUA000015997 Lee 12 May 1959 Brady, Jr., V.E.
AUA
Cynodon Dactylon (L.) Pers.
JSU108260 Bibb 22 Jul 1982 Whetstone, R. David
11657
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015988 Crenshaw 27 May 1994 Diamond, Alvin R.
9200
AUA
Cynodon Dactylon (L.) Pers.
TROY000011825 Pike 09 Nov 2000 Windham, Mark
10
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108262 Calhoun 14 Jun 1969 Landers, Kenneth E.
s.n.
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000011827 Dale 11 Oct 2000 Johnson, Stephanie
1
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0042995 Baldwin 19 Apr 2003 Larsen, Harry S.
1158
UWAL
Cynodon dactylon (L.) Pers.
TROY000011829 Dale 13 Jul 1997 Rundell, Hannelore
275
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0039749 Covington 02 Jul 2016 Diamond, Alvin R.
27535
UWAL
Cynodon dactylon (Linnaeus) Persoon
TROY000011831 Escambia 07 May 1995 Brittain, Cynthia L.
159
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0010114 Bullock 10 Sep 2011 Dykes, Bailee
416
UWAL
Cynodon dactylon (Linnaeus) Persoon
SAMF0001294 Tuscaloosa 01 Jun 1955 Robinson, Flynt
s.n.
SAMF
Cynodon dactylon (Linnaeus) Persoon
JSU108265 Cleburne 01 Oct 1996 Hruska, Susan E.
1288
JSU
Cynodon dactylon (Linnaeus) Persoon
UNA00003668 Tuscaloosa _ Jun 1955 Robinson, Flynt
s.n.
UNA
Cynodon dactylon (L.) Pers. var. dactylon
TROY000034215 Conecuh 29 Oct 2011 Diamond, Alvin R.
22669
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108268 DeKalb 05 Jul 1969 Beaty, Norma J.
s.n.
JSU
Cynodon dactylon (Linnaeus) Persoon
UWAL0056776 Marion 26 May 2023 England, Kevin J.
13652
UWAL
Cynodon dactylon (L.) Persoon
TROY000032185 Marengo 06 May 2009 England, J. Kevin
2079
TROY
Cynodon dactylon (Linnaeus) Persoon
AUA000038058 Covington 4 May 2012 Hansen, Curtis J.
5512
AUA
Cynodon Dactylon (L.) Pers.
TROY000037181 Coosa 12 Jun 2013 Diamond, Alvin R.
24235
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0008835 Marengo 06 May 2009 England, J. Kevin
2079
UWAL
Cynodon dactylon (L.) Pers.
TROY000034122 Bullock 10 Sep 2011 Dykes, Bailee
104
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0013274 Coosa 12 Jun 2013 Diamond, Alvin R.
24235
UWAL
Cynodon dactylon (Linnaeus) Persoon
UNA00003698 Tuscaloosa 28 Apr 1955 Farmer, Joe A.
s.n.
UNA
Cynodon dactylon (L.) Pers. var. dactylon
AUA000043815 Lee 5 Oct 2018 Taylor, Chris T.
CSP-557
AUA
Cynodon Dactylon (L.) Pers.
TROY000047755 Autauga 05 Oct 2018 Parker, Matthew R.
1825
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108267 Cullman 12 Aug 1974 Atkinson, Tim A.
623
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000011821 Barbour 06 Nov 2000 Green, Clint
6
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0012243 Pickens 11 Sep 2011 Keener, Brian R.
6695
UWAL
Cynodon dactylon (Linnaeus) Persoon
UNA00003639 Tuscaloosa 26 Jul 1952 Yates, Jack C.
s.n.
UNA
Cynodon dactylon (L.) Pers.
UWAL0010686 Conecuh 29 Oct 2011 Diamond, Alvin R.
22669
UWAL
Cynodon dactylon (Linnaeus) Persoon
UNA00003653 Tuscaloosa 15 Mar 1955 Mallory, Jack C.
s.n.
UNA
Cynodon dactylon (L.) Pers. var. dactylon
AUA000015992 Lee 19 Jun 1969 Miller, L.C.
031
AUA
Cynodon Dactylon (L.) Pers.
TROY000011828 Dale 18 Jul 2000 Woods, Michael
8176
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108271 Franklin 06 Jun 1975 Whetstone, R. David
4462
JSU
Cynodon dactylon (Linnaeus) Persoon
JSU108273 Jefferson 09 May 1984 Barber, J. Patrick
211
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015983 Butler 19 Jun 1988 Diamond, Alvin R.
5048
AUA
Cynodon Dactylon (L.) Pers.
UWAL0049131 Bullock 25 May 2016 Horne, Howard E.
3210
UWAL
Cynodon dactylon (Linnaeus) Persoon
TROY000046048 Montgomery 19 Jun 2010 Diamond, Alvin R.
21424
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108342 Randolph 05 Jul 1969 Herren, Tommy
s.n.
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000016002 Tallapoosa 1 May 2006 Diamond, Alvin R.
16337
AUA
Cynodon Dactylon (L.) Pers.
JSU108280 Mobile 09 Apr 1976 Wayner, Gary A.
622
JSU
Cynodon dactylon (Linnaeus) Persoon
UNA00003639 Tuscaloosa 26 Jul 1952 Yates, Jack C.
s.n.
UNA
Cynodon dactylon (L.) Pers.
TROY000011824 Coffee 28 Apr 2000 Martin, Brian H.
636
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108263 Clay 07 Jul 1999 Brodeur, Kristin R.
2043
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000026705 Tallapoosa 01 May 2006 Diamond, Alvin R.
16337
TROY
Cynodon dactylon (Linnaeus) Persoon
AUA000015987 Conecuh 27 Jun 1987 Diamond, Alvin R.
3428
AUA
Cynodon Dactylon (L.) Pers.
TROY000011833 Pike 26 Apr 1977 Whitley, Crystal D.
3
TROY
Cynodon dactylon (Linnaeus) Persoon
JSU108264 Clay 30 Jul 1994 Ballard, J. Mark
4261
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000011830 Dale 15 May 1995 Jernigan, Clay
6
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0051227 Jefferson 28 May 1984 Barber, J. Patrick
398
UWAL
Cynodon dactylon (L.) Persoon
TROY000011826 Coffee 10 Sep 1999 Martin, Brian H.
197
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0008834 Choctaw 08 May 2007 Harrell, Elizabeth A.
428
UWAL
Cynodon dactylon (L.) Pers.
UNA00003678 Tuscaloosa 16 Jun 1955 Farmer, Joe A.
s.n.
UNA
Cynodon dactylon (L.) Pers. var. dactylon
AUA000015990 Lee 12 May 1970 Williams, S.A.
147
AUA
Cynodon Dactylon (L.) Pers.
JSU156348 Etowah 14 Jun 1974 Atkinson, Tim A.
301
JSU
Cynodon dactylon (Linnaeus) Persoon
TROY000011835 Pike 21 Aug 2000 Hall, James A.
127
TROY
Cynodon dactylon (Linnaeus) Persoon
UWAL0052068 Madison 04 Oct 2020 England, J. Kevin
11327
UWAL
Cynodon dactylon (L.) Pers.
UNA00060902 Mobile 25 Dec 2002 Zebryk, Tad M.
8178
UNA
Cynodon dactylon (L.) Pers.
UNA00003653 Tuscaloosa 15 Mar 1955 Mallory, Jack C.
s.n.
UNA
Cynodon dactylon (L.) Pers. var. dactylon
UWAL0056170 Fayette 19 Jun 2023 England, Kevin J.
13756
UWAL
Cynodon dactylon (L.) Persoon
JSU108347 Talladega 26 Aug 1980 Whetstone, R. David
10299
JSU
Cynodon dactylon (Linnaeus) Persoon
UNA00003683 Mobile s.d. Mohr, Charles
s.n.
UNA
Cynodon dactylon (L.) Pers.
UNA00054651 Blount 16 Jun 1998 Keener, Brian R.
971
UNA
Cynodon dactylon (L.) Pers.
AUA000015998 Lee 15 May 1922 Simmons, W.E.
AUA
Cynodon Dactylon (L.) Pers.
JSU108344 St. Clair 13 Nov 1998 Spaulding, Daniel D.
10541
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015989 Dale 23 Jul 1989 Diamond, Alvin R.
6090
AUA
Cynodon Dactylon (L.) Pers.
JSU108348 Walker 12 Jun 1975 Whetstone, R. David
4729
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015993 Lee 16 May 1969 Sheldon, Joe
18.2
AUA
Cynodon Dactylon (L.) Pers.
TROY000044170 Covington 02 Jul 2016 Diamond, Alvin R.
27535
TROY
Cynodon dactylon (Linnaeus) Persoon
AUA000016003 Tallapoosa 18 Nov 1962 Rogers, Ken E.
AUA
Cynodon Dactylon (L.) Pers.
JSU108276 Jefferson 17 Sep 1983 Whetstone, R. David
12446
JSU
Cynodon dactylon (Linnaeus) Persoon
AUA000015995 Lee 3 May 1959 Beckert, Heino
AUA
Cynodon Dactylon (L.) Pers.
JSU108279 Marshall 21 Jul 1975 Whetstone, R. David
5189
JSU
Cynodon dactylon (Linnaeus) Persoon
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