Lithospermum tuberosum

Family:BORAGINACEAE
Species:Lithospermum tuberosum Rugel ex A. DC.
Common Name:Tuberous Gromwell; Southern Stoneseed
Habitat:Mesic hardwood forests, ravines, and bluffs.
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Perennial
Category:Vascular
USDA Symbol:LITU2
Plant Notes:Tuberous Gromwell is a native perennial in the Borage family (Boraginaceae). It can be found in all but the southernmost counties in Alabama. Tuberous Gromwell grows in mesic hardwood forests, in rocky woods, in prairie woods, in ravines, and on bluffs. Tuberous Gromwell is most common in areas underlain by limestone. It is a perennial with thickened tuberous-like roots. Stems are from 6-15 inches in height. The stems branch in the upper portions. The stems are covered in long hairs. Leaves are basal and cauline. The basal leaves form a rosette that persist until flowering. Leaves are oblanceolate to elliptic in outline, entire, and pubescent. The hairs on the leaves have a swollen base (postulate). Stem leaves are alternate and smaller than basal leaves. Flowers are produced in leafy-bracted cymes. The cymes continue to elongate as flowering and fruiting progresses. The corolla is yellowish or greenish-white in color. The corolla has a short tube and five spreading star shaped lobes. The fruit is a white, wrinkled, and stone-like mericarp.--A. Diamond
Taxonomic Notes:**
Status:Native
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyBORAGINACEAE - Borage family
Genus Lithospermum
Species Lithospermum tuberosum Rugel ex A. DC. - Tuberous Gromwell; Southern Stoneseed

Citation

Citation Lithospermum tuberosum Rugel ex A.L.P.P. de Candolle, Prodr. [A. P. de Candolle] 10: 76–77. 1846.
Basionym: **
Type: USA: FLORIDA: Gadsden Co.: Near Aspalaga,

** Not applicable or data not available.

Synonyms

No synonyms exist for this species.

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Bibb, Blount, Butler, Chambers, Chilton, Clarke, Colbert, Conecuh, Dallas, Etowah, Fayette, Franklin, Jackson, Jefferson, Lauderdale, Lawrence, Lowndes, Madison, Marengo, Marshall, Monroe, Montgomery, Morgan, Shelby, St. Clair, Sumter, Talladega, Tallapoosa, Tuscaloosa, Wilcox

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AMAL, AUA, JSU, TROY, UNA, USAM, UWAL

Range of years during which specimens were collected: 1962 - 1982

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
JSU114907 Jackson 25 Apr 1973 Whetstone, R. David
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00056603 Bibb 01 May 1953 Brown, Jack S.
s.n.
UNA
Lithospermum tuberosum Rugel ex DC.
USAM000003070 Monroe 10 Apr 1970 Wofford, B. Eugene
10416
USAM
Lithospermum tuberosum Rugel ex A.P. de Candolle
AMAL00005982 Bibb 23 Apr 1953 Taxonomy class-Spring 1953
s.n.
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
ALNHS00008552 Madison 05 May 1993 Oberholster, Chris
CO#654
ALNHS
Lithospermum tuberosum Rugel
JSU114906 Jackson 14 Apr 1973 Kral, Robert
49586
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
TROY000029569 Butler 19 Mar 2009 Diamond, Alvin R.
20535
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114893 Bibb 13 May 1983 Whetstone, R. David
13427
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA000042896 Montgomery 10 Apr 2020 Birkhead, Roger
AUA
Lithospermum Tuberosum Ugel.
UNA00012880 Tuscaloosa 1 Apr _ Smith, Eugene A.
s.n.
UNA
Lithospermum tuberosum Rugel ex DC.
AUA_ACC_38479 Madison 8 Apr 1972 Dooley, M.K.
58
AUA
Lithospermum Tuberosum Ugel.
TROY000032378 Butler 17 Apr 2011 Diamond, Alvin R.
21909
TROY
Lithospermum tuberosum Rugel ex A.P. del Candolle
UWAL0007986 Jackson 12 Apr 2008 Keener, Brian R.
4208
UWAL
Lithospermum tuberosum Rugel ex DC.
ALNHS00007478 Dallas 13 APR 2022 Barger, T. Wayne
SP#5738
ALNHS
Lithospermum tuberosum Rugel
AMAL00020066 Dallas 15 Apr 2010 Barger, T. Wayne
47
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00058687 Lawrence 17 Apr 2005 Keener, Brian R.
2626
UNA
Lithospermum tuberosum Rugel ex DC.
UWAL0006054 Monroe 10 Apr 1983 Kral, Robert
69871
UWAL
Lithospermum tuberosum Rugel ex DC
JSU114890 Bibb 22 Jul 1982 Whetstone, R. David
11748
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0034672 Lowndes 04 Oct 1994 Kral, Robert
84276
UWAL
Lithospermum tuberosum Rugel ex DC
JSU114915 Morgan 29 Mar 1997 Whetstone, R. David
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00039478 Wilcox 16 Apr 1980 Haynes, Robert R.
7720
UNA
Lithospermum tuberosum Rugel ex DC.
UNA00039469 Dallas 09 Apr 1982 Gunn, Scott C.
397
UNA
Lithospermum tuberosum Rugel ex DC.
JSU114899 Fayette 8 Jun 1999 Haynes, Robert R.
9975
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_38821 Autauga 7 Sep 1982 Gunn, Scott C.
1194
AUA
Lithospermum Tuberosum Ugel.
UNA00066014 Lawrence 06 May 1999 Haynes, Robert R.
9947
UNA
Lithospermum tuberosum Rugel ex DC.
JSU114909 Madison 22 Sep 1995 Threlkeld, Steven J.
82
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00039477 Bibb 11 Apr 1979 Wiersema, John H.
523
UNA
Lithospermum tuberosum Rugel ex DC.
UWAL0007991 Sumter 19 Mar 2007 Keener, Brian R.
3421
UWAL
Lithospermum tuberosum Rugel ex DC.
UNA00039472 Bibb _ _ 1953 Taxonomy class-Spring 1953
s.n.
UNA
Lithospermum tuberosum Rugel ex DC.
AMAL00020035 Monroe 06 Apr 2011 Barger, T. Wayne
65
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00039468 Autauga 07 Sep 1982 Gunn, Scott C.
1194
UNA
Lithospermum tuberosum Rugel ex DC.
AUA_ACC_4006 Clarke 19 Mar 1955 Davis, Donald E.
AUA
Lithospermum Tuberosum Ugel.
UNA00039475 Tuscaloosa 25 Mar 1962 Deramus, Rebecca
78
UNA
Lithospermum tuberosum Rugel ex DC.
TROY000032194 Marengo 17 Apr 2009 England, J. Kevin
1972
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114902 Jackson 24 Oct 1992 Ballard, J. Mark
781
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0007987 Jackson 15 Apr 2007 Keener, Brian R.
3495
UWAL
Lithospermum tuberosum Rugel ex DC.
JSU114905 Jackson 4 May 1994 Spaulding, Daniel D.
6128
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_25674 Franklin 16 Apr 1965 Kral, Robert
23565
AUA
Lithospermum Tuberosum Ugel.
AUA_ACC_37518 Marshall 23 Aug 1981 O'Halloran, K.
438
AUA
Lithospermum Tuberosum Ugel.
JSU114891 Autauga 7 Sep 1982 Gunn, Scott C.
1194
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_38815 Wilcox 16 Apr 1982 Gunn, Scott C.
505
AUA
Lithospermum Tuberosum Ugel.
UWAL0019091 Franklin 14 Apr 1998 Kral, Robert
87285
UWAL
Lithospermum tuberosum Rugel ex DC
TROY000044911 Autauga 02 Mar 2017 Parker, Matthew R.
376
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0032373 Franklin 14 Apr 2016 England, J. Kevin
6210
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
AMAL00013858 Lawrence 17 Apr 2005 Keener, Brian R.
2626
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00039481 Monroe 10 Mar 1968 McDaniel, Sidney
10352
UNA
Lithospermum tuberosum Rugel ex DC.
AUA_ACC_33013 Bibb 8 Jul 1976 Sessler, Ann H.
193
AUA
Lithospermum Tuberosum Ugel.
TROY000016767 Jackson 31 Mar 1973 Whetstone, R. David
s.n.
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114913 Marshall 12 Jun 1993 Whetstone, R. David
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0046847 Autauga 02 Mar 2017 Parker, Matthew R.
376
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA000044003 Madison 6 Mar 2020 Hansen, Curtis J.
7828
AUA
Lithospermum Tuberosum Ugel.
TROY000032495 Dallas 12 May 2011 Diamond, Alvin R.
22087
TROY
Lithospermum tuberosum Rugel ex A.P. del Candolle
JSU114898 Etowah 20 Aug 1994 Hodge, Lesley M.
3086
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AMAL00005985 Madison 05 May 1996 Threlkeld, Steven J.
229
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
TROY000016787 Chilton 03 May 2005 Diamond, Alvin R.
15286
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114896 Etowah 28 Apr 1972 Whetstone, R. David
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
TROY000036589 Sumter 11 Mar 2012 Keener, Brian R.
6934
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114901 Franklin 14 May 1993 Whetstone, R. David
16368
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA000039198 Morgan 19 May 1934 Wolf, Wolfgang
AUA
Lithospermum Tuberosum Ugel.
TROY000029789 Jackson 13 May 2009 Barger, T. Wayne
121
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_19257 Chambers 11 Jul 1969 Moore, Harold D.
257-69
AUA
Lithospermum Tuberosum Ugel.
TROY000016786 Autauga 29 Jul 1989 Diamond, Alvin R.
6177
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114900 Fayette 18 Mar 1979 Whetstone, R. David
9423
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0011683 Colbert 17 Jun 2013 Keener, Brian R.
7918
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
TROY000038256 Butler 02 Apr 2013 Diamond, Alvin R.
23890
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_72070 Jefferson 11 Apr 2012 Barger, T. Wayne
519
AUA
Lithospermum Tuberosum Ugel.
JSU114894 Blount 17 Aug 1997 Ballard, J. Mark
7368
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0007989 Morgan 04 Apr 2009 England, J. Kevin
1836
UWAL
Lithospermum tuberosum Rugel ex DC.
JSU114911 Marshall 23 Apr 1993 Spaulding, Daniel D.
3500
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_57214 Jackson 14 Apr 1973 Kral, Robert
49586
AUA
Lithospermum Tuberosum Ugel.
JSU114916 Morgan 28 May 1975 Whetstone, R. David
4111
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
TROY000045628 Autauga 15 Sep 2016 Parker, Matthew R.
75
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_38822 Autauga 12 May 1982 Gunn, Scott C.
530
AUA
Lithospermum Tuberosum Ugel.
UWAL0018543 Lowndes 06 Apr 1993 Kral, Robert
81996
UWAL
Lithospermum tuberosum Rugel ex DC
TROY000036108 Madison 27 Mar 2012 Keener, Brian R.
6959
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0007988 Marengo 17 Apr 2009 England, J. Kevin
1972
UWAL
Lithospermum tuberosum Rugel ex DC.
AMAL00021666 Lowndes 16 Apr 2014 Keener, Brian R.
8339
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0033890 Fayette 30 Mar 2016 Keener, Brian R.
9423
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
TROY000016769 Sumter 19 Mar 2007 Keener, Brian R.
3421
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_35024 Jefferson 7 Jul 1977 Sessler, Ann H.
1697
AUA
Lithospermum Tuberosum Ugel.
TROY000030939 Clarke 19 Mar 1955 Davis, Donald E.
s.n.
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114917 Morgan 10 Jun 1976 Whetstone, R. David
8427
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0044866 Colbert 20 Apr 2019 England, J. Kevin
10137
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114912 Marshall 13 Sep 1991 Spaulding, Daniel D.
927
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0015579 Lowndes 16 Apr 2014 Keener, Brian R.
8339
UWAL
Lithospermum tuberosum Rugel ex DC.
JSU114914 Marshall 29 Mar 1976 Whetstone, R. David
7794
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0009726 Sumter 11 Mar 2012 Keener, Brian R.
6934
UWAL
Lithospermum tuberosum Rugel ex DC.
UWAL0052958 Bibb 24 Jul 2019 Keener, Brian R.
11483
UWAL
Lithospermum tuberosum Rugel ex. A DC.
JSU114910 Madison 1 Apr 1973 Gurley, Bob
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_38820 Lowndes 6 Sep 1982 Gunn, Scott C.
1125
AUA
Lithospermum Tuberosum Ugel.
UNA00039471 Tuscaloosa 11 Apr 1969 Benton Jr., Cole L.B.
s.n.
UNA
Lithospermum tuberosum Rugel ex DC.
UNA00039480 St. Clair 28 Apr 1979 Davenport, Lawrence J.
1084
UNA
Lithospermum tuberosum Rugel ex DC.
AUA_ACC_41056 Bibb 13 May 1983 Whetstone, R.David
13427
AUA
Lithospermum Tuberosum Ugel.
UNA00039479 Madison 13 Apr 1980 Meigs, Russell A.
230
UNA
Lithospermum tuberosum Rugel ex DC.
JSU114908 Lauderdale 14 Apr 1969 D. W.
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0005507 Dallas 12 May 2011 Diamond, Alvin R.
22088
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
AMAL00020114 Lauderdale 28 Jun 2010 Holt, Brian D.
377
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0046500 Autauga 15 Sep 2016 Parker, Matthew R.
75
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00012881 s.d. Smith, Eugene A.
s.n.
UNA
Lithospermum tuberosum Rugel ex DC.
UWAL0007992 Tuscaloosa 30 Mar 2009 Keener, Brian R.
5124
UWAL
Lithospermum tuberosum Rugel ex DC.
JSU114918 St. Clair 22 May 1972 Whetstone, R. David
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0044765 Jackson 02 Jun 2018 England, J. Kevin
9187
UWAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
JSU114920 Tuscaloosa 3 Mar 1979 Whetstone, R. David
9340
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00039474 Lawrence 26 Apr 1977 Haynes, Robert R.
6114
UNA
Lithospermum tuberosum Rugel ex DC.
JSU114919 Talladega 14 May 1982 Bussey, Melanie G.
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00039470 Lowndes 06 Sep 1982 Gunn, Scott C.
1125
UNA
Lithospermum tuberosum Rugel ex DC.
JSU114895 Etowah 12 Jun 1994 Hodge, Lesley M.
2360
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_38818 Dallas 9 Apr 1982 Gunn, Scott C.
397
AUA
Lithospermum Tuberosum Ugel.
UNA00053299 Fayette 08 Jun 1999 Haynes, Robert R.
9975
UNA
Lithospermum tuberosum Rugel ex DC.
AUA_ACC_63323 Monroe 4 Apr 2003 Conti, Andrea
AUA
Lithospermum Tuberosum Ugel.
ALNHS00000475 Dallas 15 Apr 2010 Barger, T. Wayne
OC-47
ALNHS
Lithospermum tuberosum Rugel
UWAL0009752 Madison 27 Mar 2012 Keener, Brian R.
6959
UWAL
Lithospermum tuberosum Rugel ex DC.
UNA00039482 Tuscaloosa 13 Apr 1966 Thomas, Joab L.
796
UNA
Lithospermum tuberosum Rugel ex DC.
AMAL00014670 Tallapoosa 13 Jun 2006 Barger, T. Wayne
203
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UNA00053852 Blount 13 Jun 1997 Keener, Brian R.
201
UNA
Lithospermum tuberosum Rugel ex DC.
AUA_ACC_33562 Bibb 12 Apr 1977 Sessler, Ann H.
923
AUA
Lithospermum Tuberosum Ugel.
TROY000034969 Butler 22 Apr 2012 Diamond, Alvin R.
22956
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0012379 Butler 02 Apr 2013 Diamond, Alvin R.
23890
UWAL
Lithospermum tuberosum Rugel ex A.P. Candolle
JSU114897 Etowah 13 Apr 1996 Hodge-Spaulding, Lesley M.
4441
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
AUA_ACC_43579 Wilcox 6 Apr 1985 Diamond, Alvin R.
313
AUA
Lithospermum Tuberosum Ugel.
JSU114903 Jackson 28 Apr 1994 Whetstone, R. David
s.n.
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
TROY000016768 Jackson 15 Apr 2007 Keener, Brian R.
3495
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
AMAL00005983 Etowah 14 Apr 1995 Spaulding, Daniel D.
7900
AMAL
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0006032 Jackson 07 Jun 1982 Kral, Robert
68684
UWAL
Lithospermum tuberosum Rugel ex DC
TROY000014733 St. Clair 26 May 2008 Keener, Brian R.
4486
TROY
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0007990 St. Clair 26 May 2008 Keener, Brian R.
4486
UWAL
Lithospermum tuberosum Rugel ex DC.
JSU114892 Bibb 18 May 1984 Whetstone, R. David
13945
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0052557 Bibb 16 Mar 2019 Keener, Brian R.
11032
UWAL
Lithospermum tuberosum Rugel ex A. DC.
TROY000040789 Colbert 17 Jun 2013 Keener, Brian R.
7918
TROY
Lithospermum tuberosum Rugel ex A. P. de Candolle
AUA_ACC_49483 Conecuh 30 Jan 1988 Diamond, Alvin R.
4102
AUA
Lithospermum Tuberosum Ugel.
UWAL0054171 Shelby 11 Apr 2020 Spaulding, Daniel D.
16894
UWAL
Lithospermum tuberosum Rugel ex DC.
JSU114904 Jackson 1 May 1998 Brodeur, Kristin R.
627
JSU
Lithospermum tuberosum Rugel ex A.P. de Candolle
UWAL0031458 Colbert 23 Apr 2011 England, J. Kevin
2815
UWAL
Lithospermum tuberosum Rugel ex AP de Candolle
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