Modiola caroliniana

Family:MALVACEAE
Species:Modiola caroliniana (L.) G. Don
Common Name:Carolina Bristle Mallow; Cheeses; Bristly Mallow
Habitat:**
Associated Ecological Communities:**
Growth Habit:Herb
Duration:Perennial
Category:Vascular
USDA Symbol:MOCA
Plant Notes:Carolina Bristle Mallow is a native perennial in the Hibiscus family (Malvaceae). It can be found throughout Alabama. Carolina Bristle Mallow occurs in lawns, on roadsides, along railroads, on shell mounds, on dunes, in pine woods, in open hardwood forests, in fields, in prairies, and on rock outcrops. It is a perennial form a tuberous rootstock. Stems radiate out from the crown. Stems are usually prostrate and root at the nodes. Occasional plants have a more up-right growth form. The stems are pubescent. Leaves are alternate, petiolate, pubescent, and vary in outline from deltoid to reniform. The leaves are often deeply 3-7 palmately lobed with toothed or incised margins. The stipules are persistent and leaf-like. Flowers are produce solitarily from the leaf axils. There are 3 narrow lanceolate segments just below the calyx (the epicalyx). There are 5 sepals and 5 orange-red petals. The stamens are united into a column that is yellow in color. The fruit is a schizocarp consisting of 20 reniform mericarps in a single whorl. The mericarps are black in color and have bristles and 2 spines. Each mericarp is 2 seeded. Carolina Bristle Mallow is often weedy and occurs in a wide variety of habitats. It is drought and salt tolerant.--A. Diamond
Taxonomic Notes:**
Status:Native, FACU+ (NWI)
References:**
Specimen: View specimen details in the Alabama Herbarium Consortium Specimen Database

** Not applicable or data not available.

Classification

FamilyMALVACEAE - Mallow family
Genus Modiola
Species Modiola caroliniana (L.) G. Don - Carolina Bristle Mallow; Cheeses; Bristly Mallow

Citation

Citation Modiola caroliniana (Linnaeus) G. Don, Gen. Hist. 1: 466. 1831.
Basionym: Malva caroliniana L. 1753.
Type: "Carolina", without data (lectotype: LINN 870.15). Lectotypified by Paul & Nayar, Fasc. Fl. India 19: 183. 1988.

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Abutilodes caroliniana Abutilodes caroliniana (Linnaeus) Kuntze, Revis. Gen. Pl. 1: 65. 1891.Malva caroliniana Linnaeus 1753. 
Anoda strictiflora Anoda strictiflora Steudel, Flora 39: 437. 1856. CHILE: Without data, Bertero 406 (holotype: ?; isotype: P).
Malva caroliniana Malva caroliniana Linnaeus, Sp. Pl. 688. 1753. "Carolina", without data (lectotype: LINN 870.15). Lectotypified by Paul & Nayar, Fasc. Fl. India 19: 183. 1988.
Malva prostrata Malva prostrata Cavanilles, Diss. 2: 59, t. 16(3). 1786. URUGUAY: Near Montevideo, 1767, Commerson s.n. (holotype: P).
Malva urticifolia Malva urticifolia Kunth, in Humboldt et al., Nov. Gen. Sp. 5: 276. 1822. COLOMBIA: Cundinamarca: Santa Fe de Bogota, s.d., Humboldt & Bonpland s.n. (holotype: P).
Modanthos caroliniana Modanthos caroliniana (Linnaeus) Alefeld, Oesterr. Bot. Z. 13: 12. 1863.Malva caroliniana Linnaeus 1753. 
Modanthos prostrata Modanthos prostrata (Cavanilles) Alefeld, Oesterr. Bot. Z. 13: 12. 1763.Malva prostrata Cavanilles 1786. 
Modanthos reptans Modanthos reptans (A. Saint-Hilaire) Alefeld, Oesterr. Bot. Z. 13: 12. 1863.Modiola reptans A. Saint-Hilaire 1827. 
Modanthos urticifolia Modanthos urticifolia (Kunth) Alefeld, Oesterr. Bot. Z. 13: 12. 1863.Malva urticifolia Kunth 1822. 
Modiola multifida Modiola multifida Moench, Methodus 620. 1794, nom. illegit.Malva caroliniana Linnaeus 1753. 
Modiola prostrata Modiola prostrata (Cavanilles) A. Saint-Hilaire, Fl. Bras. Merid. 1: 211. 1827.Malva prostrata Cavanilles 1786. 
Modiola reptans Modiola reptans A. Saint-Hilaire, Fl. Bras. Merid. 1: 212, t. 43. 1827. URUGUAY: Cerro San Miguel, "ad limites provinciarum Cisplatanae et Río Grande do Sul", s.d., Saint-Hilaire s.n. (holotype: P).
Modiola urticifolia Modiola urticifolia (Kunth) G. Don, Gen. Hist. 1: 466. 1831.Malva urticifolia Kunth 1822. 

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Baldwin, Barbour, Bullock, Butler, Calhoun, Cherokee, Chilton, Choctaw, Clarke, Clay, Cleburne, Coffee, Colbert, Conecuh, Crenshaw, Escambia, Etowah, Fayette, Geneva, Jackson, Jefferson, Lauderdale, Lawrence, Lee, Marengo, Marshall, Mobile, Montgomery, Morgan, Pickens, Pike, Russell, Shelby, St. Clair, Sumter, Talladega, Tallapoosa, Tuscaloosa, Washington

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AUA, JSU, SAMF, TROY, UNA, UNAF, UWAL

Range of years during which specimens were collected: 1949 - 2019

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
AUA_ACC_3432 Lee 2 May 1949 Cunningham, Hugh B.
AUA
Modiola Caroliniana (L.) G. Don
UNA00067198 Butler 18 Apr 2007 Diamond, Alvin R.
17401
UNA
Modiola caroliniana (L.) G. Don
UNA00007416 Lawrence 09 May 1987 Haynes, Robert R.
9104
UNA
Modiola caroliniana (L.) G. Don
JSU132204 Calhoun 16 Apr 1969 Clonts, Jerry A.
s.n.
JSU
Modiola caroliniana (Linnaeus) G. Don
TROY000007005 Crenshaw 30 Apr 2000 Diamond, Alvin R.
11664
TROY
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_42078 Conecuh 9 May 1986 Diamond, Alvin R.
2464
AUA
Modiola Caroliniana (L.) G. Don
UWAL0006731 Marengo 17 Apr 2009 England, J. Kevin
1984
UWAL
Modiola caroliniana (L.) G. Don
JSU132215 Shelby 12 Oct 1996 Spaulding, Daniel D.
9429
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_43985 Montgomery 14 May 1986 Diamond, Alvin R.
2504
AUA
Modiola Caroliniana (L.) G. Don
UNA00010987 Tuscaloosa 29 Aug 1960 Harper, Roland M.
4463
UNA
Modiola caroliniana (L.) G. Don
AUA_ACC_3430 Lee 17 Apr 1949 Snow, Samuel P.
AUA
Modiola Caroliniana (L.) G. Don
UWAL0018382 Baldwin 29 Mar 1986 Kral, Robert
73210
UWAL
Modiola caroliniana (L.) G.Don
AUA_ACC_3428 Lee 10 May 1952 Shell
AUA
Modiola Caroliniana (L.) G. Don
TROY000021403 Chilton 10 Apr 2005 Diamond, Alvin R.
15088
TROY
Modiola caroliniana (Linnaeus) G. Don
UWAL0050787 Lee 16 Apr 1973 Villadsen, Martha
159
UWAL
Modiola caroliniana (L.) G. Don
UNAF0001743 Colbert 05/00/1973 Reynolds, B.
s.n.
UNAF
Modiola caroliniana (Linnaeus) G. Don
UNA00010903 Mobile 22 Apr 1950 Crawford, Lloyd C.
951
UNA
Modiola caroliniana (L.) G. Don
AUA_ACC_46489 Montgomery 10 May 1975 Veal, Fred
140
AUA
Modiola Caroliniana (L.) G. Don
JSU132207 Clay 30 Jul 1994 Ballard, J. Mark
4248
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA000036675 Escambia 12 Sep 2011 Hansen, Curtis J.
5265
AUA
Modiola Caroliniana (L.) G. Don
UNA00010952 Pickens 04 Jul 1986 Horn, Charles N.
1124
UNA
Modiola caroliniana (L.) G. Don
JSU142223 Clarke 21 Mar 2012 Horne, Howard E.
SWS-99
JSU
Modiola caroliniana (Linnaeus) G. Don
JSU132209 Etowah 14 Sep 1994 Hodge, Lesley M.
3830
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_36077 Russell 4 May 1973 Scruggs, Debra A.
89
AUA
Modiola Caroliniana (L.) G. Don
JSU132199 Calhoun 7 May 1986 Lawler, Carol L.
408
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_3431 Lee 28 Apr 1949 Dean, Dorothy
AUA
Modiola Caroliniana (L.) G. Don
UWAL0056651 Fayette 29 Mar 2019 England, Kevin J.
10047
UWAL
Modiola caroliniana (Linnaeus) G.Don
UNA00022253 Marengo 11 Apr 1969 Thomas, Joab L.
1960 1/2
UNA
Modiola caroliniana (L.) G. Don
JSU132202 Calhoun 6 Apr 1967 Collier, Gary
s.n.
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_22457 Lee 10 May 1970 Knotts, Carl R.
187
AUA
Modiola Caroliniana (L.) G. Don
UWAL0014683 Bullock 22 Mar 2012 Dykes, Bailee
412
UWAL
Modiola caroliniana (Linnaeus) G. Don
UNAF0001742 Colbert 10 May 1973 Dalrymple, Larry
s.n.
UNAF
Modiola caroliniana (Linnaeus) G. Don
SAMF0007740 Jefferson 30 Mar 2017 Parrish, Preston
8
SAMF
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_1510 Lee 18 Jul 1962 Gramlich, Jim
AUA
Modiola Caroliniana (L.) G. Don
TROY000036702 Bullock 22 Mar 2012 Dykes, Bailee
412
TROY
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_22198 Lee 5 May 1970 Sullivan, Terry K.
84
AUA
Modiola Caroliniana (L.) G. Don
UWAL0050584 Sumter 06 Apr 2021 Ryan, Emma D.
71
UWAL
Modiola caroliniana (L.) G. Don
AUA_ACC_64539 Chilton 10 Apr 2005 Diamond, Alvin R.
15088
AUA
Modiola Caroliniana (L.) G. Don
ALNHS00004390 Lee 04 Apr 2016 Taylor, Chris T.
CSP-276
ALNHS
Modiola caroliniana (L.) G. Don
UWAL0002170 Chilton 13 Apr 2008 Spaulding, Daniel D.
12830
UWAL
Modiola caroliniana (L.) G. Don
TROY000007004 Conecuh 07 Apr 2001 Diamond, Alvin R.
12338
TROY
Modiola caroliniana (Linnaeus) G. Don
JSU132214 Marshall 22 Apr 1992 Spaulding, Daniel D.
1354
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_55977 Clarke 1 Jun 1972 Kral, Robert
47069
AUA
Modiola Caroliniana (L.) G. Don
UWAL0043395 Baldwin 10 Apr 2006 Larsen, Harry S.
1498
UWAL
Modiola caroliniana (L.) G. Don
UNAF0001744 Lee 19 Apr 1962 Landers, Kenneth E.
s.n.
UNAF
Modiola caroliniana (Linnaeus) G. Don
SAMF0007639 Shelby 18 Apr 2021 Thompson, Madelyn
57
SAMF
Modiola caroliniana (Linnaeus) G. Don
UWAL0006732 St. Clair 06 May 2008 Keener, Brian R.
4397
UWAL
Modiola caroliniana (L.) G. Don
TROY000028152 St. Clair 06 May 2008 Keener, Brian R.
4397
TROY
Modiola caroliniana (Linnaeus) G. Don
UWAL0049725 Lauderdale 28 Apr 2008 Webb, David H.
6739
UWAL
Modiola caroliniana (L.) G. Don
JSU132201 Calhoun 4 Jun 1993 Ballard, J. Mark
2737
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_14933 Lee 19 May 1969 Harmon, Crystal
157
AUA
Modiola Caroliniana (L.) G. Don
TROY000007001 Barbour 01 May 1997 Adams, David L., Jr
7
TROY
Modiola caroliniana (Linnaeus) G. Don
UWAL0006734 Sumter 10 Apr 1993 Ray, Stephan
s.n.
UWAL
Modiola caroliniana (L.) G. Don
TROY000007002 Butler 18 Apr 2007 Diamond, Alvin R.
17410
TROY
Modiola caroliniana (Linnaeus) G. Don
UNA00010913 Tuscaloosa 02 Apr 1949 Crawford, Lloyd C.
s.n.
UNA
Modiola caroliniana (L.) G. Don
AUA_ACC_67600 Cleburne 14 Apr 2006 Tuggle, Andrea
11
AUA
Modiola Caroliniana (L.) G. Don
JSU132216 Talladega 29 Mar 1997 Ballard, J. Mark
7155
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_32574 Russell 7 Apr 1974 Lancaster, D.S.
141
AUA
Modiola Caroliniana (L.) G. Don
UNA00010992 Mobile 30 Apr 1966 Deramus, Rebecca
D885
UNA
Modiola caroliniana (L.) G. Don
UWAL0043396 Baldwin 07 May 2004 Larsen, Harry S.
1373
UWAL
Modiola caroliniana (L.) G. Don
JSU132208 Colbert 24 May 1974 Atkinson, Tim A.
235
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_2637 Tallapoosa 16 May 1965 Brown, Jimmy E.
AUA
Modiola Caroliniana (L.) G. Don
JSU132200 Calhoun 2 May 1987 Lawler, Carol L.
1584
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_61127 Choctaw 6 Jun 1994 Crouch, Virginia E.
997B
AUA
Modiola Caroliniana (L.) G. Don
UNA00066820 Butler 18 Apr 2007 Diamond, Alvin R.
17401
UNA
Modiola caroliniana (L.) G. Don
UNA00010982 Tuscaloosa 10 May 1959 Edwards, Jackie
s.n.
UNA
Modiola caroliniana (L.) G. Don
UWAL0037498 Bullock 06 Apr 2016 Soehren, Eric C.
216
UWAL
Modiola caroliniana (L.) G. Don
TROY000007007 Pike 27 Apr 1993 Coleman, Dennis
6
TROY
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_49156 Pike 5 May 1988 Diamond, Alvin R.
4647
AUA
Modiola Caroliniana (L.) G. Don
UWAL0017272 Lawrence 07 Jul 2007 England, J. Kevin
497
UWAL
Modiola caroliniana (Linnaeus) G.Don
JSU132213 Marshall 10 May 1992 Spaulding, Daniel D.
1593
JSU
Modiola caroliniana (Linnaeus) G. Don
JSU132206 Cherokee 3 Nov 1996 Whetstone, R. David
s.n.
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_61128 Choctaw 24 Apr 1994 Crouch, Virginia E.
906
AUA
Modiola Caroliniana (L.) G. Don
UWAL0006733 Marengo 01 May 1993 Harris, J.W.
s.n.
UWAL
Modiola caroliniana (L.) G. Don
UWAL0006730 Butler 18 Apr 2007 Diamond, Alvin R.
17401
UWAL
Modiola caroliniana (L.) G. Don
JSU132212 Lee 11 May 1966 Berry, Robert
s.n.
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_49105 Pike 24 Jun 1988 Diamond, Alvin R.
5067
AUA
Modiola Caroliniana (L.) G. Don
UNA00010908 Mobile 23 Apr 1950 Crawford, Lloyd C.
1019
UNA
Modiola caroliniana (L.) G. Don
AUA_ACC_3429 Lee 13 May 1952 Adkins, Jr., T.R.
AUA
Modiola Caroliniana (L.) G. Don
AUA_ACC_22199 Washington 11 Apr 1970 Williams, Ann
43
AUA
Modiola Caroliniana (L.) G. Don
UNA00010972 Mobile 27 Mar 1965 Deramus, Rebecca
D 302
UNA
Modiola caroliniana (L.) G. Don
UWAL0032416 Morgan 23 Apr 2016 England, J. Kevin
6246
UWAL
Modiola caroliniana (Linnaeus) G. Don
UNA00010957 Shelby 09 May 1981 Horn, Charles N.
256
UNA
Modiola caroliniana (L.) G. Don
JSU132203 Calhoun 6 Feb 1993 Spaulding, Daniel D.
3342
JSU
Modiola caroliniana (Linnaeus) G. Don
ALNHS00001390 Mobile 20 Mar 2012 Barger, T. Wayne
SP#726
ALNHS
Modiola caroliniana (L.) G. Don
AUA_ACC_11394 Lee 20 Apr 1966 Searcy, James W.
AUA
Modiola Caroliniana (L.) G. Don
TROY000007006 Escambia 29 Apr 1995 Brittain, Cynthia L.
102
TROY
Modiola caroliniana (Linnaeus) G. Don
UWAL0044685 Lauderdale 28 May 2019 England, J. Kevin
10272
UWAL
Modiola caroliniana (Linnaeus) G. Don
JSU132211 Lee 10 Apr 1964 Bozarth, Gene A.
s.n.
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_19642 Lee 29 Apr 1969 Moore, Harold D.
139-69
AUA
Modiola Caroliniana (L.) G. Don
UNA00010947 Mobile _ May 1878 Mohr, Charles
s.n.
UNA
Modiola caroliniana (L.) G. Don
AUA_ACC_3427 Lee 27 Apr 1949 Arline, Marvin H.
AUA
Modiola Caroliniana (L.) G. Don
UWAL0005518 Conecuh 04 Jun 2011 Diamond, Alvin R.
22174
UWAL
Modiola caroliniana (Linnaeus) G. Don
TROY000007003 Coffee 03 May 2000 Martin, Brian H.
676
TROY
Modiola caroliniana (Linnaeus) G. Don
JSU132210 Jackson 26 May 1999 Brodeur, Kristin R.
1867
JSU
Modiola caroliniana (Linnaeus) G. Don
TROY000032745 Conecuh 04 Jun 2011 Diamond, Alvin R.
22174
TROY
Modiola caroliniana (Linnaeus) G. Don
JSU132205 Calhoun 30 Apr 1968 Stuart, Martha
s.n.
JSU
Modiola caroliniana (Linnaeus) G. Don
AUA_ACC_3426 Lee 4 May 1949 Cope, Will A.
AUA
Modiola Caroliniana (L.) G. Don
TROY000042395 Chilton 13 Apr 2008 Spaulding, Daniel D.
12830
TROY
Modiola caroliniana (Linnaeus) G. Don
AUA000038436 Morgan 23 Apr 2016 England, J. Kevin
6246
AUA
Modiola Caroliniana (L.) G. Don
TROY000045209 Geneva 23 Mar 2017 Byrd, Cameron
874
TROY
Modiola caroliniana (L.) G. Don
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